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smut fungus
biol. 黑粉菌属 (Ustilago (Pers.) Rouss.，黑粉菌科); 黑粉菌 (Ustilago sp.，指黑穗病，黑粉病的病菌); 黑粉菌科 (Ustilaginaceae)
ecol. 黑粉病菌; 黑穗病菌
life.sc. 黑粉菌

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